The only differences I can see in the CR2 between bone_1 and bone_2  that seem to relate to this are highlighted in yellow on the comparison screenshots below - bone_1 on the left, bone_2 on the right.

Firstly, the twistY, jointX and jointZ are in a different position and order in the channels section and are named in a different manner

 

Secondly, the twistY, jointX and jointZ of bone_1 all have two extra parameters - flipped and primaryParm.

I've checked Kuroyume's unofficial CR2 spec but 2004 was a long time ago and it doesn't really give me any clues, except that the order of jointX and jointZ is important

While I wait for somebody who knows what they're talking about I may just try editing a copy of the CR2 to make the bone_1 joint parameters similar to the bone_2 ones... what's the worst that can happen ? ;o)

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The 3Dcheapskate (also available in DAZ and HiveWire3D flavours) occasionally posts sensible stuff. Usually by accident.
And it usually uses Poser 11, with units set to inches. Except when it's using Poser 6 or PP2014, or when its units are set to PNU.